Wild food plants used by the Tibetans of Gongba Valley (Zhouqu county, Gansu, China)
© Kang et al.; licensee BioMed Central Ltd. 2014
Received: 23 December 2013
Accepted: 27 January 2014
Published: 6 February 2014
The ethnobotany of Tibetans is a seriously under-studied topic. The aim of the study was to investigate knowledge and use of wild food plants in a valley inhabited by Tibetans in the Gannan Tibetan Autonomous Region.
The field research was carried out in a wooded mountain valley in 9 neighbouring villages the Zhouqu (Brugchu) county, and comprised 17 interviews with single informants and 14 group interviews, involving 122 people altogether.
We recorded the use of 81 species of vascular plants from 41 families. Fruits formed the largest category, with 42 species, larger than the wild greens category, with 36 species. We also recorded the culinary use of 5 species of edible flowers, 7 species with underground edible organs and 5 taxa of fungi. On average, 16.2 edible taxa were listed per interview (median – 16). Green vegetables formed the largest category of wild foods (mean – 8.7 species, median – 9 species), but fruits were listed nearly as frequently (mean – 6.9, median – 6). Other categories were rarely mentioned: flowers (mean – 0.2, median – 0), underground edible parts (mean – 0.3, median – 0) and mushrooms (mean – 1.5, – median 1).
Wild vegetables are usually boiled and/or fried and served as side-dishes (cai). They are often lacto-fermented. Wild fruits are mainly collected by children and eaten raw, they are not stored for further use. The most widely used wild vegetables are: Eleuterococcus spp., Pteridium aquilinum, Helwingia japonica, Aralia chinensis, Allium victorialis, Pteridium aquilinum, Ixeris chinensis, Thlaspi arvense and Chenopodium album. The culinary use of Caltha palustris as a green vegetable is very interesting. In its raw state, marsh marigold is a toxic plant, due to the presence of protoanemonin. In this area it is dried or lactofermented before use. The most commonly eaten fruits are: Pyrus xerophila, Prunus salicina, Berchemia sinica, Rubus spp. and Eleagnus umbellata.
The number of wild taxa eaten in the studied valley is relatively large compared to most studies from around the world. However, compared to the northern slope of the Qinling, in Shaanxi, the list is considerably shorter, in spite of the similar methodology applied and similar research effort involved.
KeywordsWild edible plants Brugchu Ethnobotany Ethnomycology Caltha palustris Lactic fermentation Lacto-fermented
Wild food plants still function as an important part of human nutrition in many parts of the globe. Although nowadays they usually provide only a minor proportion of daily calories, they are important sources of vitamins, microelements and other health supporting chemicals. They have always enabled human survival in times of crises [1–5]. The diversity of wild food plants consumed depends mainly on which species are available in sufficient quantity in the surrounding vegetation, the socioeconomic status of the population and cultural attitudes towards different categories of wild food. China is a very interesting arena for studies on wild food use traditions, as in all three above-mentioned aspects there are conditions that support the wide use of plants . It contains an immense diversity of habitats and possesses numerous biodiversity hotspots. Some of its populations in remote rural areas still have a relatively low economic status, which binds people to traditional subsistence agriculture and exploitation of natural resources. Moreover, the rural population exploits different categories of wild food, including numerous species of wild vegetables, and is in itself diverse, consisting of many ethnic minorities, which differ in their plant utilization habits.
The ethnobotany research in China, concerning wild food plants, has so far been concentrated mainly on Yunnan [7–12] and Inner Mongolia [13, 14], also as far as wild food is concerned. Although some papers on the use of wild food plants have been published from other parts of China (e.g. Sichuan [15, 16], Hunan  etc.), some provinces are seriously under-studied ethnobotanically. An example of such a place is the province of Gansu, for which we found only one small article on wild vegetables, and only for the easternmost part of the province inhabited by Han Chinese . Gansu is an ethnic mosaic of a few ethnic groups: Han Chinese, Hui and Tibetans . In spite of the tremendous cultural diversity of different Tibetan ethnic groups and their diverse plant use, there is relatively little ethnobotanical literature devoted to the Tibetan people in China [12, 20–22]. There is a particularly scarce ethnobotanical literature concerning the use of wild food plants by Tibetans, also in other provinces, with only one major article on the wild food uses of the Tibetans in Shangri-La (NW Yunnan) .
Although Tibetans are classified in China as one minority (族藏, Zangzu), they consist of a mosaic of several Tibetan languages, dialects and ways of life, with six main groups/dialects, out of which three occur in China classified as Ü-Tsang, Kham and Amdo . The linguistic situation on the Gansu – Sichuan border is little studied, and it is one of the regions in which ever newer dialects are described (e.g. ). Some linguists distinguish over twenty separate distinct languages in the Tibetan group of languages . As Nicolas Tournandre puts it: “there are 220 ‘Tibetan dialects’ derived from Old Tibetan and nowadays spread across 5 countries: China, India, Bhutan, Nepal and Pakistan which may be classed within 25 dialect groups, i.e. groups which do not allow mutual intelligibility. (…) Thus if we set aside the notion of standardization, I believe it would be more appropriate to speak of 25 languages derived from Old Tibetan. This is not only a terminological issue but it gives an entirely different perception of the range of variation. When we refer to 25 languages, we make clear that we are dealing with a family comparable in size to the Romance family which has 19 groups of dialects”.
To fill the gap in the ethnobotanical exploration of the northwestern part of China we aimed at documenting the use of wild food plants in one Tibetan valley in SW Gansu, on the edge of the Qinling Mountains and the Tibetan Plateau. As we have been, since 2011, carrying out similar research in other parts of the Qinling, inhabited by Han Chinese, one of the research questions also compared the pattern of wild food plant use with our previous studies [26–28].
Tibetans in Gannan constitute a very diverse collection of subsistence economies and speak a variety of dialects. In the northern part of the territory, in the grasslands, animal herders predominate. The Diebu (Tewo) Tibetans, although living among forests, live mainly from animal husbandry, whereas in the southern part of the region we studied, in some forested valleys, plant cultivation is the main source of subsistence.
The Chagang Township has an area of 157 km2. The average annual rainfall is 680 mm (mainly in summer months), the annual average temperature is 9.1°C, and the frost-free period lasts 220 days. The average annual net income is 1376 Yuan. The total population is 3624, of which farmers account for 3197. There are Tibetan, Han and Hui in the township, but the Tibetan population accounts for 83% of the total population .
Wuping township is located upstream from the Chagang township. It has an area of (429 km2). The annual average temperature ranges between 6°C and 12°C, the frost-free period lasts 160 to 200 days. It consists of 10 administrative villages, including 32 small settlements, with a total population of 6962. There are 6240 farmers, including 2889 Tibetans, who constitute 41.5% of the total population .
The studied villages are purely Tibetan and usually set at the top of a hill, with a few dozen houses, all very close to each other, e.g. Jiaorao Village, with 83 households and 483 inhabitants, Gudang – 62 houses, 300 people, Gaerli – over 500 people and Hadie – 116 houses, 618 inhabitants . We studied nine villages altogether, in Chagangzhen township (插岗镇): Jiaorao (角绕), Jiaoerli (噶儿里), Caopo (草坡), Qinggangpo (青岗坡), Chagang (插岗村), Gudang (古当), Wubie (吾别); in Wupingzhen township (武坪镇): Hadie (哈迭) and Wuping (武坪). The altitude ranged from 1782 to 2140 m a.s.l.
The studied population consists of subsistence farmers cultivating barley, potatoes, maize, flax, buckwheat, broad bean, pea, rape-seed and some fruit trees, e.g. apple, pear, walnuts and Sichuan pepper (Zanthoxylum bungeanum). More rarely fungi (mainly Auricularia and Grifola umbellata) and medicinal herbs (Gastrodia, Bupleurum chinense, Codonopsis) are cultivated.
The field research was conducted in August 2013 using the Rapid Rural Appraisal approach [30, 31], and included 31 freelisting interviews (17 interviews with single informants and 14 group interviews), which altogether involved 122 people (70 men and 52 women).The mean age of the participants was 50 (median 48, aged from 12 to 85; 52 women and 70 men). For the seventeen people (11 men and 6 women) who were interviewed separately the mean age was 53, median 51, youngest interviewee – 23, the oldest one – 75).
The research was carried out following the code of ethics of the American Anthropological Association  and the International Society of Ethnobiology Code of Ethics . Oral prior informed consent was acquired. Statistics were calculated using the open access statistical package PAST [37, 38].
Results and discussion
Twenty most frequently mentioned wild food plants in the freelisting questionnaire
Frequency (N = 31)
Eleuterococcus giraldii & E. leucorrhisus
Ferns in general (‘zhala’)
Rubus mesogaeus & R. thibetanus
Young aerial parts
Young aerial parts, roots
Young aerial parts
Wild food plants collected by at least two informants
Local Chinese name
Local Chinese name (pinyin)
Local Tibetan name (broad transcription of pronunciation according to IPA)$
Actinidia tetramera Maxim.
Amaranthus retroflexus L.*
Chenopodium album L.
Hui Hui Cai, Hui Tiao
Acroglochin persicarioides Moq.*
Fu Hui Tiao
tieniliәmә [literally ‘sheep Chenopodium’]
Allium victorialis L.
Luo Er Jiu
老葱, 野蒜, 野葱
Lao Cong, Ye Suan, Ye Cong
Rhus verniciflua Stokes
Carum carvi L.
Ye Hui Xiang
Eleutherococcus giraldii (Harms) T.Nakai ,
Wu Zhua Cai
Eleutherococcus leucorrhizus Oliv.
Wu Zhua Cai
Aralia chinensis L.
Mu Ian Tou
Artemisia subdigitata Mattf.*
Niu Wei Hao
Ixeris chinensis Nakai.
Ku Ma Cai
Prenanthes macrophylla Franch.*
Shan Jian Zi
Tong Tong Cai
Sinacalia tangutica (Maxim.) B.Nord.**
paibɔ, piaba, paibie
Sonchus oleraceus L.
Tong Tong Cai
wu: [mainly], (ng)wu:, muwawu:, muwawu:
Taraxacum cf mongolicum Han.-Mzt
Pu Gong Ying
Brassica campestris L.
Cardamine macrophylla Willd.
Shi Gen Cai
Thlaspi arvense L.
Ku Gen Cai
Adenophora stricta Miq.
Lonicera standishii Carr.
Qiao Pi Zi
(k)alaɨala, asa, , kai/gai, ɕiese(ng)
Lonicera tragophylla Hemsl. ex Forb. & Hemsl.
Yu Yu Hua
mudesɨbe, nɨmenu, :
Triosteum pinnatifidum Maxim.
kelu, keɨü, kalɔilɔ, alaɨala
Triosteum himalayanum Wall.
Viburnum betulifolium Batalin
Viburnum glomeratum Maxim.
Cephalotaxus sinensis (Rehder & E.H.Wilson) H.L.Li
Shui Bai Zi
Cornus japonica Thunb.
Corylus sieboldiana Blume
Corylus heterophylla Fisch. ex Besser
Corylus ferox Wall.
Pteridium aquilinum (L.) Kuhn var. latiusculum
Elaeagnus umbellata Thunb.
Hippophae rhamnoides L.
Sophora alopecuroides L.
Helwingia japonica (Thunb.) F.
Ye Li Kai Hua
Juglans cathayensis Dode
Ye He Tao
nina, dienanu, dienane = walnut flɔwers[die = walnut tree]
Caryopteris divaricata Maxim.
Mentha haplocalyx Briq.*
Stachys affinis Bunge
Decaisnea fargesii Franch
Gui Zhi Tou
Akebia trifoliata (Thunb.) Koidz.
Da Hua, Bai He
Malva verticillata L.
(ng)nagu, nɨnagu, ɨamuda
Toona sinensis (A.Juss.) M.Roem.
Morus australis Poir.
Ye Sang Zi, Ye Sang Shen
Matteucia intermedia C.Chr.
Matteucia struthiopteris (L.) Tod.
Pinus armandii Franch.
Huang Song, Su Mu Tou
tɔm(u)gu, tɔm(u)gə tɔr(ə), tɔʐɨ
Plantago depressa Willd.*
Persicaria alata (Buch.-Ham.) Nakai*
Caltha palustris L.
Huang Hua Cai
Berchemia sinica C.K.Schneid.
Ti Ge Er
saü, sawi, saɨu
Rhamnus rosthornii E.Pritz. ex Diels
Crataegus wilsonii Sarg.
Crataegus kansuensis E.H.Wilson
Fragaria pentaphylla Losinsk.
Maddenia hypoleuca Koehne
Malus baccata (L.) Borkh.
Shan Ding Zi
Neillia sinensis Oliv.
mahadәʐә, wahadәʐә, wahadәr, mahuɔtu
Prunus davidiana Franch.
Ye Mao Tao
ti:, ting, kuaʐә, kuebu, kuɔʐuɔ, kuumɔ
Prunus salicina Lindl
Ku Li Zi, Ye Li Zi
Prunus tomentosa Thunb.
Pyrus xerophila T.T.Yu
Ye Suan Li
ɕieʐɨ, ɕiedɔ, ɕiɔ:, ɕialә(ng)
Rosa brunonii Lindl.
Rubus amabilis Focke
Hong Mao Zi
ʐәna, ʐɨna, ʐɨna mɨngʐɨ, ʐɨna ɕiʐu
Rubus mesogaeus Focke
Hei Mao Zi
Rubus thibetanus Franch
Hei Mao Zi
ʐәna, ʐɨna, duheni, ɕiehele, wunihebie
Rubus xanthocarpus Bureau & Franch.
Huang Mao Zi
Sorbus xanthoneura Rehder
Huang Mai Hua Qiu
Zanthoxylum bungeanum Maxim.
Jiao Ya, Hua Jiao
Populus davidiana Dode
Yang Shu Hua
Buckleya henryi Diels
Rodgersia aesculifolia Batalin**
Gui Deng Qing
Schisandra sphenanthera Rehder & E.H.Wilson
Wu Wei Zi
Staphylea holocarpa Hemsl.
Liang Zi Jian
Pseudocyclosorus subochthoides (Ching) Ching
Urtica fissa E.Pritz. ex Diels*
sana [mainly], [rarely]
Vitis piasezkii Maxim.
Ye Pu Tao
Wild mushrooms collected for food mentioned by at least two informants
Local Chinese name
Local Chinese name (pinyin)
Local Tibetan name (pronunciation according to IPA)
Grifola umbellata (Pers.) Pilát
Zhu Ling Hua
Morchella spp. (mainly M. angusticeps Peck)
Yang Du Jun
Auricularia spp. (mainly A. auricula-judae (Bull.) Quél.)
Ye Mu Er
“a red capped mushroom”
Hong Mo Gu
On average, 16.2 edible taxa were listed per interview (median – 16). The most listed category of wild foods was green vegetables (mean – 8.7 species, median – 9 species), but fruits were listed nearly as frequently (mean – 6.9, median – 6). Other category lists were very short: flowers (mean – 0.2, median – 0), underground edible parts (mean – 0.3, median – 0) and mushrooms (mean – 1.5, – median 1).
In group interviews we obtained only slightly more species per list (e.g. fruits – 6.4 species in individual ones, 6.9 in group ones; green vegetables – 8.3 and 9.2 respectively). The difference between individual and group interviews is small, probably due to the fact that some of the best informants were interviewed separately.
Wild vegetables are usually boiled and/or fried and served as side-dishes (cai). Wild fruits are collected mainly by children and eaten raw, they are not stored for further use. Mushrooms are usually fried. During the 1959–61 famine Sinacalia tangutica tubers (one of the commonest plants in the roadsides) were widely used as emergency food and they are well remembered, although no one eats them nowadays.
The most widely used wild vegetables are: Eleuterococcus spp., ferns - called collectively zhala (pinyin pronunciation), Helwingia japonica, Aralia chinensis, Allium victorialis, Pteridium aquilinum, Ixeris chinensis, Thlaspi arvense and Chenopodium album (Table 1). The first four of these taxa (Eleuterococcus spp., ferns, Helwingia japonica and Aralia chinensis) are also traded throughout the county and are mentioned even in the Zhouqu county annals as important sources of income for the locals .
It is interesting that some wild greens, which are very common species in the valley and are eaten in other parts of China, e.g. wild amaranths Amaranthus spp. and nettles Urtica spp. [6, 26, 27], are almost entirely ignored, regarded as inedible by most of the population, and eaten by very few individuals.
The most commonly eaten fruits are: wild pears Pyrus xerophila, wild plums Prunus salicina, Berchemia sinica, blackberries Rubus spp. (black fruited) and Eleagnus umbellata. The large number species of fruits reported in this study, compared to the central part of the Qinling Mts. [26, 27], probably stems from the different landscape. Here, there is an abundance of natural hedgerows growing on steep banks dividing the field terraces, providing large masses of edible fruits near homesteads.
the local flora is relatively poorer here (the rainfall is lower, forest understory is poorer and the forest vegetation is more transformed by humans)
the local population is less interested in using all the potentially edible wild vegetables, for example the extremely abundant Amaranthus and Urtica are not utilized by the majority of the population.
In our previous papers [26, 27] from northern Qinling we noted that local people seem to value forest wild greens over the ruderal taxa. This is even more visible in this part of the Qinling, where the gathering efforts are nearly exclusively oriented towards forest greens. Local people venture six km hikes up steep slopes in order to collect ferns, marsh marigolds Caltha palustris and bittercress Cardamine macrophylla, and do not bother to collect annual wild greens growing around their homesteads!
The culinary use of Caltha palustris is very interesting. In its raw state marsh marigold is a highly toxic plant, due to the presence of protoanemonin, and has not been eaten recently. We only have historical reports about its use in northern Europe, mainly from Estonia , and from the United States (as potherb – , or flower buds as caper substitutes – ). Some plant dictionaries have also recorded the edibility of Caltha leaves and roots, e.g. Kluk’s Dykcyonarz Roślinny from eighteenth century Poland . Protoanemonin is broken down by drying and long cooking [43, 44]. In the study area, the plant is either dried and then used like other wild greens, or it is lacto-fermented after the initial blanching. The dried specimens of marsh marigolds we ate had no bitterness even prior to cooking. Further studies are needed to establish if the local population of marsh marigolds is a low-protoanemonin form easier to prepare as food than the European Caltha. For example, one of the authors’ (ŁŁ) unpublished experiments show that C. palustris from Poland needs one to two hours of cooking to get rid of the bitterness caused by protoanemonin. Some species of the related genus Ranunculus have been eaten in Europe after only short cooking, as they are not as bitter as European Caltha, e.g. R. repens L. in Belarus and Poland , R. neapolitanus Ten. in Croatia , Ranunculus muricatus L. in Herzegovina  and R. ficaria L. in Slovakia , Ukraine , Romania and Hungary  and Italy .
It is interesting that the list of wild edible fruits is slightly longer than that of wild greens, as the reverse was usually the case in wild food studies in East Asia. The structure of the edible species list (similar number of wild greens and fruits) is very similar to that recorded among the Tibetans in Shangri-La  where 80 wild vegetable species and 78 edible fruit species were recorded. It is also more reminiscent of other regions of the world such as Africa [51, 52] and India [53, 54], where wild greens are used widely but not as many species as in the Qinling Mountains and other regions of East Asia, where the local biodiversity enables a choice from many species of wild vegetables.
Similarly to the Shaanxi part of the Qinling mountains, the number of fungi taxa collected is low. There were only four taxa mentioned by more than four informants, whereas as many as 153 species of edible mushrooms were identified in the area . The two most commonly collected taxa (Grifola and Morchella) do not have original Tibetan names, but names borrowed from Chinese, which suggests that it was the Chinese who introduced their use. In contrast to the mushrooms, there is high consensus on wild vegetable and fruit names and all of them possess local Tibetan names.
The high level of bilinguality as far as plant names are concerned is worth emphasizing. Although bi- and trilinguality is very common in cultural edge areas (Zhouqu county has been a Tibetan-Chinese ethnic borderland for centuries), a nearly full bilinguality concerning plant names is a much rarer phenomenon. When we asked our informants why they know nearly all the local Chinese names of edible plants (although they speak only Tibetan in their villages) they explained that they need them when they talk about plants with people from the neighbouring Han villages. This is a proof that fortunately wild edible plants still constitute an important topic of conversation for the villagers and they maintain a high cultural status.
The number of wild taxa eaten in the studied valley is relatively large compared to most studies of edible plants from throughout the world. However, compared to the northern slope of the Qinling, the list is shorter, in spite of the similar methodology applied and similar research effort involved. The culinary use of the toxic Caltha palustris is worth emphasizing.
Many thanks to the inhabitants of the studied villages for their willingness in sharing information on the use of the species. The program was financially supported by the Forestry Research Foundation for the Public Service Industry of China (2009,04004) and by the University of Rzeszów (Institute of Biotechnology and Basic Sciences, as well as a special grant from the rector of the university). Many thanks to prof. Xiaosi Guo for the identification of ferns.
- Maurizio A: Die Geschichte unserer Pflanzennahrung: von den Urzeiten bis zur Gegenwart. 1927, Berlin: PareyGoogle Scholar
- Johns T: With bitter herbs they shall eat it: chemical ecology and the origins of human diet and medicine. 1990, Tucson: University of Arizona PressGoogle Scholar
- Bharucha Z, Pretty J: The roles and values of wild foods in agricultural systems. Phil Trans R Soc B. 2010, 365: 2913-2926. 10.1098/rstb.2010.0123.PubMed CentralView ArticlePubMedGoogle Scholar
- Delang C: Not just minor forest products: the economic rationale for the consumption of wild food plants by subsistence farmers. Ecol Econ. 2006, 59: 64-73. 10.1016/j.ecolecon.2005.10.006.View ArticleGoogle Scholar
- Turner NJ, Łuczaj ŁJ, Migliorini P, Pieroni A, Dreon AL, Sacchetti LE, Paoletti MG: Edible and tended wild plants, traditional ecological knowledge and agroecology. Cr Rev Plant Sci. 2011, 30 (1–2): 198-225.View ArticleGoogle Scholar
- Hu SY: Food Plants of China. 2005, Hongkong: The Chinese University PressGoogle Scholar
- Long C-L, Li H, Ouyang Z, Yang X, Li Q, Trangmar B: Strategies for agrobiodiversity conservation and promotion: a case from Yunnan, China. Biodivers Conserv. 2003, 12 (6): 1145-1156. 10.1023/A:1023085922265.View ArticleGoogle Scholar
- Xu Y, Tao G, Liu H, Yan K, Dao X: Wild vegetable resources and market survey in Xishuangbanna, Southwest China. Econ Bot. 2004, 58 (4): 647-667. 10.1663/0013-0001(2004)058[0647:WVRAMS]2.0.CO;2.View ArticleGoogle Scholar
- Zhang XP, Wu JL, Li Y, Liu F, Wang JQ: Investigation on species resources and utilization of wild vegetable in Nabanhe watershed nature reserve, Xishuangbanna. J South Forestry Coll. 2004, 24: 21-24.Google Scholar
- Ghorbani A, Langenberger G, Sauerborn J: A comparison of the wild food plant use knowledge of ethnic minorities in Naban River Watershed National Nature Reserve, Yunnan, SW China. J Ethnobiol Ethnomed. 2012, 8: 17-10.1186/1746-4269-8-17.PubMed CentralView ArticlePubMedGoogle Scholar
- Ghorbani A, Langenberger G, Liu J-X, Wehner S, Sauerborn J: Diversity of medicinal and food plants as non-timber forest products in Naban river Watershed National Nature Reserve (China): implications for livelihood improvement and biodiversity conservation. Econ Bot. 2012, 66: 178-191. 10.1007/s12231-012-9188-1.View ArticleGoogle Scholar
- Ju Y, Zhuo J, Liu B, Long C: Eating from the wild: diversity of wild edible plants used by Tibetans in Shangri-la region, Yunnan, China. J Ethnobiol Ethnomed. 2013, 9 (1): 28-10.1186/1746-4269-9-28.PubMed CentralView ArticlePubMedGoogle Scholar
- Khasbagan , Huai HY, Pei SJ: Wild plants in the diet of Arhorchin Mongol herdsmen in Inner Mongolia. Econ Bot. 2000, 54 (4): 528-536. 10.1007/BF02866550.View ArticleGoogle Scholar
- Wujisguleng W, Khasbagen K: An integrated assessment of wild vegetable resources in Inner Mongolian Autonomous Region, China. J Ethnobiol Ethnomed. 2010, 6: 34-10.1186/1746-4269-6-34.PubMed CentralView ArticlePubMedGoogle Scholar
- Weckerle CS, Huber FK, Yongping Y, Weibang S: Plant knowledge of the Shuhi in the Hengduan mountains, Southwest China. Econ Bot. 2006, 60 (1): 3-23. 10.1663/0013-0001(2006)60[3:PKOTSI]2.0.CO;2.View ArticleGoogle Scholar
- Huber FK, Ineichen R, Yang K, Weckerle CS: Livelihood and conservation aspects of non-wood forest product collection in the Shaxi Valley, Southwest China. Econ Bot. 2010, 64 (3): 189-204. 10.1007/s12231-010-9126-z.View ArticleGoogle Scholar
- Zou X, Huang F, Hao L, Zhao J, Mao H, Zhang J, Ren S: The socio-economic importance of wild vegetable resources and their conservation: a case study from China. Kew Bull. 2010, 65: 577-582. 10.1007/s12225-010-9239-7.View ArticleGoogle Scholar
- Wang L, Wang R: Research on the resources of wild vegetable plants in Tianshui of Gansu Province [甘肃天水野生蔬菜植物资源研究]. Forest By-Product and Speciality in China. 2013, 122 (1): 76-77.Google Scholar
- Hansen MH: Frontier people: Han settlers in minority areas of China. 2007, Vancouver: UBC PressGoogle Scholar
- Boesi A: Plant categories and types in Tibetan materia medica. Tibet J. 2005, 30 (4/1): 65-Google Scholar
- Boesi A: Plant knowledge among Tibetan populations. Memorie della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano. 2005, 33 (1): 33-48.Google Scholar
- Liu Y, Dao Z, Yang C, Liu Y, Long C: Medicinal plants used by Tibetans in Shangri-la, Yunnan, China. J Ethnobiol Ethnomed. 2009, 5: 15-10.1186/1746-4269-5-15.PubMed CentralView ArticlePubMedGoogle Scholar
- DeLancey S: Lhasa Dialect. Sino-Tibetan Languages (Vol. 3). Edited by: Thurgood G, LaPolla RJ. 2003, London: Psychology Press, 270-288.Google Scholar
- Sun JT-S: Phonological profile of Zhongu: a new Tibetan dialect of Northern Sichuan. Lang Ling. 2003, 4 (4): 769-836.Google Scholar
- Tournadre N: Arguments against the Concept of ‘Conjunct’/‘Disjunct’ in Tibetan. Chomolangma, Demawend und Kasbek. Festschrift für Roland Bielmeier zu seinem 65. Geburtstag. Vol. 1. Edited by: Huber B, Volkart M, Widmer M, Schwieger P. 2008, 281-308.Google Scholar
- Kang Y, Łuczaj Ł, Yes S, Zhang S, Kang J: Wild food plants and wild edible fungi of Heihe valley (Qinling Mountains, Shaanxi, central China): herbophilia and indifference to fruits and mushrooms. Acta Soc Bot Pol. 2012, 81 (4): 405-413. 10.5586/asbp.2012.044.View ArticleGoogle Scholar
- Kang Y, Łuczaj Ł, Kang J, Zhang S: Wild food plants and wild edible fungi in two valleys of the Qinling Mountains (Shaanxi, central China). J Ethnobiol Ethnomed. 2013, 9 (1): 26-10.1186/1746-4269-9-26.PubMed CentralView ArticlePubMedGoogle Scholar
- Kang Y, Łuczaj Ł, Ye S: The highly toxic Aconitum carmichaelii Debeaux as a root vegetable in the Qinling Mountains (Shaanxi, China). Genet Resour Crop Evol. 2012, 59: 1569-1575. 10.1007/s10722-012-9853-3.View ArticleGoogle Scholar
- The Editorial Committee of Gansu Zhouqu Local Annals: Zhouqu County Annals. 2010, Beijing: Fangzhi Press, [甘肃舟曲县地方志编篡委员会,舟曲县志,方志出版社:北京,2010]Google Scholar
- Croll EJ: Research methodologies appropriate to rapid appraisal: a Chinese experience. IDS Bull. 1984, 15 (1): 51-56. 10.1111/j.1759-5436.1984.mp15001008.x.View ArticleGoogle Scholar
- Long CL, Wang JR: Participatory rural appraisal: An introduction to principle, methodology and application. 1996, Kunming: Yunnan Science and Technology PressGoogle Scholar
- Sutrop U: List task and a cognitive salience index. Field Methods. 2001, 3 (13): 263-276.View ArticleGoogle Scholar
- Quinlan M: Considerations for collecting freelists in the field: examples from ethnobotany. Field Methods. 2005, 17 (3): 219-234. 10.1177/1525822X05277460.View ArticleGoogle Scholar
- Guo Q, Gao Z, Song X, Li Y, Fu Z, Wang Y, He H, Tang Y: Macrofungi flora distribution and faunal study in Bailongjiang-Taohe forest region (白龙江、洮河林区大型真菌资源及分布). J Northwest Forestry University. 2010, 25 (3): 126-136.Google Scholar
- American Anthropological Association Code of Ethics. [http://www.aaanet.org/issues/policy-advocacy/upload/AAA-Ethics-Code-2009.pdf]
- International Society of Ethnobiology Code of Ethics (with 2008 additions). [http://ethnobiology.net/code-of-ethics/]
- Hammer O, Harper DAT, Ryan PD: PAST: Paleontological Statistics software package for education and data analysis. Paleontol Electron. 2001, 4 (1): 9-Google Scholar
- PAST version 2.17b. [http://folk.uio.no/ohammer/past]
- Kalle R, Sõukand R: Historical ethnobotanical review of wild edible plants of Estonia (1770 s–1960s). Acta Soc Bot Pol. 2012, 81 (4): 271-281. 10.5586/asbp.2012.033.View ArticleGoogle Scholar
- Gray A: Manual of the Botany of the Northern United States. 1848, Boston and Cambridge: James Munroe & Co.Google Scholar
- Porcher FP: Resources of the Southern Fields and Forests. Medical , Economical and Agricultural; being also a Medical Botany of the Southern States; with practical information on the useful properties of the trees, plants, and shrubs. 1869, Charleston: Walker, Evans and CogswellGoogle Scholar
- Kluk K: Dykcyonarz roślinny. Tom I. 1786, Warszawa: Drukarnia Xięży PiarówGoogle Scholar
- Gmelin L: Fortsetzung des Handbuchs der Organischen Chemie. Part 4. 1866, Heidelberg: Carl Winter’s Universitat-BuchhandlungsGoogle Scholar
- Berger A, Wachter H: Hunnius Pharmazeutisches Wörterbuch (in German). 1998, Berlin: Walter de Gruyter Verlag, 8Google Scholar
- Łuczaj Ł, Köhler P, Pirożnikow E, Graniszewska M, Pieroni A, Gervasi T: Wild edible plants of Belarus: from Rostafiński’s questionnaire of 1883 to the present. J Ethnobiol Ethnomed. 2013, 9: 21-10.1186/1746-4269-9-21.PubMed CentralView ArticlePubMedGoogle Scholar
- Łuczaj Ł, Zovko Končić M, Miličević T, Dolina K, Pandža M: Wild vegetable mixes sold in the markets of Dalmatia (Southern Croatia). J Ethnobiol Ethnomed. 2013, 9: 1-10.1186/1746-4269-9-1.View ArticleGoogle Scholar
- Łuczaj Ł: Ethnobotanical review of wild edible plants of Slovakia. Acta Soc Bot Pol. 2012, 81 (4): 245-255. 10.5586/asbp.2012.030.View ArticleGoogle Scholar
- Łuczaj Ł: Brzozowy sok, “czeremsza” i zielony barszcz – ankieta etnobotaniczna wśród botaników ukraińskich. Birch sap, ramsons and green borsch – an ethnobotanical survey among Ukrainian botanists. Etnobiologia Polska. 2012, 2: 15-22.Google Scholar
- Dénes A, Papp N, Babai D, Czúcz B, Molnár Z: Wild plants used for food by Hungarian ethnic groups living in the Carpathian Basin. Acta Soc Bot Pol. 2012, 81 (4): 381-396. 10.5586/asbp.2012.040.View ArticleGoogle Scholar
- Pieroni A: Gathered wild food plants in the upper valley of the Serchio River (Garfagnana), Central Italy. Econ Bot. 1999, 53 (3): 327-341. 10.1007/BF02866645.View ArticleGoogle Scholar
- Addis G, Urga K, Dikasso D: Ethnobotanical study of edible wild plants in some selected districts of Ethiopia. Hum Ecol. 2005, 33 (1): 83-118. 10.1007/s10745-005-1656-0.View ArticleGoogle Scholar
- Johns T, Kokwaro JO: Food plants of the Luo of Siaya district, Kenya. Econ Bot. 1991, 45 (1): 103-113. 10.1007/BF02860055.View ArticleGoogle Scholar
- Sundriyal M, Sundriyal RC: Wild edible plants of the Sikkim Himalaya: marketing, value addition and implications for management. Econ Bot. 2004, 58 (2): 300-315. 10.1663/0013-0001(2004)058[0300:WEPOTS]2.0.CO;2.View ArticleGoogle Scholar
- Rajasab AH, Isaq M: Documentation of folk knowledge on edible wild plants of north Karnataka. Indian J Tradit Know. 2004, 3 (4): 419-429.Google Scholar
This article is published under license to BioMed Central Ltd. This is an Open Access article distributed under the terms of the Creative Commons Attribution License (http://creativecommons.org/licenses/by/2.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly credited. The Creative Commons Public Domain Dedication waiver (http://creativecommons.org/publicdomain/zero/1.0/) applies to the data made available in this article, unless otherwise stated.