An ethnobotanical study of the less known wild edible figs (genus Ficus) native to Xishuangbanna, Southwest China
© Shi et al.; licensee BioMed Central Ltd. 2014
Received: 8 July 2014
Accepted: 17 September 2014
Published: 24 September 2014
The genus Ficus, collectively known as figs, is a key component of tropical forests and is well known for its ethnobotanical importance. In recent decades an increasing number of studies have shown the indigenous knowledge about wild edible Ficus species and their culinary or medicinal value. However, rather little is known about the role of these species in rural livelihoods, because of both species and cultural diversity.
In this study we 1) collected the species and ethnic names of wild edible Ficus exploited by four cultural groups in Xishuangbanna, Southwest China, and 2) recorded the collection activities and modes of consumption through semi-structured interviews, 3) investigated the resource management by a statistical survey of their field distribution and cultivation, and 4) compared and estimated the usage intensities by the grading method.
The young leaves, leaf buds and young or ripe syconia of 13 Ficus species or varieties are traditionally consumed. All the species had fixed and usually food-related ethnic names. All four cultural groups are experienced in the collection and use of edible Ficus species as vegetables, fruits or beverages, with the surplus sold for cash income. Different cultural groups use the Ficus species at different intensities because of differences in availability, forest dependency and cultural factors. Both the mountain and basin villagers make an effort to realize sustainable collection and meet their own and market needs by resource management in situ or cultivation.
In comparison with reports from other parts of the world, ethnic groups in Xishuangbanna exploited more edible Ficus species for young leaves or leaf buds. Most of the edible species undergo a gradient of management intensities following a gradient of manipulation from simple field gathering to ex situ cultivation. This study contributes to our understanding of the origins and diffusion of the knowledge of perception, application and managing a group of particular plant species, and how the local culture, economic and geographical factors influence the process.
The genus Ficus consists of over 800 species of trees, shrubs, vines and epiphytes in the family Moraceae, which have a wide distribution and multiple uses in most tropical and subtropical regions throughout the world. They are traditionally used as medicine or food plants, ornamental trees, religious plants, lacca hosts, fodder, fuel wood, hedges or enclosures. The importance of Ficus as a global spiritual and material resource for humans has been well-documented [1, 2].
Some Ficus species are reported to be among the oldest human food sources. Fossil evidence suggests that the common fig (Ficus carica L.) has been cultivated for over 11,000 years, possibly predating cereal grains , and thousands of cultivars of this species have been developed worldwide. The biblical sycamore fig (F. sycomorus L.) is another old human food source and one of the largest fruit producers. It was identified as a key food source for Pliocene hominids along the Baragoi River in Nachola and Baragoi, in northern Kenya . Nowadays the most well-known species is F. carica, which produces the commercial common fig (fig is also the general name of the “fruit” of the genus). Specifically, the common fig is the syconium of F. carica var. domestica, which is functionally a female individual of F. carica. However, it was only in recent decades that it became known how many more culinary uses the genus Ficus can provide beyond their nutritious raw syconia. A growing number of studies have noted the significance of the edible Ficus species as vegetables [5–9], for tea or beverage preparation [10–12], for jelly and jam production  or for medicinal purposes [14–17].
Our previous ethnobotanical survey conducted in southwest Yunnan, China, found that Ficus was the most frequently consumed genus (with 8 species) of the 220 plant genera which contain species used as wild vegetables . Ficus was also the most frequently used genus (7 species) among the 82 genera containing wild fruits in the region . Specialized studies of these wild edible Ficus species are needed to clarify their ethnobotany, including vernacular names, modes of consumption, availability and management, and possible multiple uses. Since most of the species identified in these surveys are native to the Xishuangbanna Dai Autonomous Prefecture, a cultural and biodiversity hotspot in southwest Yunnan, China, a research project was initiated to estimate the current and historic usage of edible Ficus species in Xishuangbanna, not only for their economic benefits, but also for their cultural richness and their roles in the long history of rural livelihoods.
Information on the ethnic villages sampled in Xishuangbanna, SW China
Main income sources
Rubber trees, wild collection
Cash crops (bean and corn), livestock, wild collection and hunting
Rubber trees, farmland rent, part – time jobs
Cash crops (Amomum villosum, banana, etc.), livestock, wild collection and hunting
Ethnobotanical data collection
Semi-structured interviews were conducted with 10–27 key informants in each village in March–October 2012, for details about the wild edible Ficus species, including their local nomenclatures, collection techniques and modes of consumption. The number of interviewees selected in each village mainly depended on the integrity and uniformity of information we obtained. Only those who could give details independently and according to their own experience, or could give a practical demonstration, were counted as key informants. Interviewees were first shown a group of sample species (photographs or fresh specimens) that were expected to be used. Species mentioned by informants that were not included in the samples were given special attention as new records. The Yao villagers living in Dazhai are monolingual, with only the village school students and staff able to speak Mandarin, so our interviews were conducted in the school with Yao students and teachers. Dai, Hani and Jinuo interviews were conducted in the villagers’ homes, farming plots or collection sites using Mandarin or their own language, with the help of local field assistants.
Field investigations were designed to assess the availability of edible fig resources, including their field distribution and home cultivation. Since the mountain villages (Bakaxiaozhai, Dazhai and Nanbang) do not have private homegardens, they consume only wild Ficus. We chose Nanbang as a representative mountain village to assess the distribution of the fig individuals they use. Six key informants were asked to guide field travel in the area surrounding their village in March 2012. Ficus individuals in dietary use and within one-hour walking distance (along twisting narrow footpaths in five directions) from the village site were recorded, including their species names, the numbers of plants, and their diameters at breast height (DBH). We chose “one-hour walking distance” to include most plots where the daily collection activities happen and the distance was accessible for all of our local guides. The Dai, who settle in valleys, basins or on river banks at lower elevations, are the major culture group in XSBN and have developed diverse homegardens. Most Ficus parts they eat come from these or from around their courtyards. We visited 107 households in Manlun village in May–June 2012 to collect data regarding the species, numbers of plants, and DBH of all edible Ficus species cultivated.
Observations of trading in Ficus products were conducted in three local markets related to the selected villages. They are the farm products market of 1) Menglun township, 2.2 km to Manlun and 5.7 km to Bakaxiaozhai, 2) Mengla township, 31 km to Nanbang, and 3) Jinghong city, which assemble farm products and wild collections from multiple ethnic groups and wide areas. All the villages can access their markets by motor vehicles. We selected Menglun market for long-term (November 2013 to April 2014) monitoring to collect basic data (species, price, sources, etc.) and determine the amounts sold and peak season, because the market has the most diverse Ficus products and modest size. We did market survey in Mengla and Jinghong during peak season to assess the universality of the trading activities. Only part of the result is shown in this study. More details on the amounts sold and their sources will be included in another publication because it needed additional investigations.
Based on the usage intensity classification for wild fruits of Chen et al. , we defined five levels of usage intensity according to our work described above. The five levels are coded as 1) –, no usage record; 2) +, consumed occasionally or just by children and hunters; 3) + +, frequently consumed and often gathered regularly during harvest season; 4) + + +, frequently consumed and the surplus may be sold in the local market or processed for out-of-season use, sources protected; 5) + + + +, cultivated in homegardens for consumption and sale.
All voucher specimens collected in our field work were deposited in the herbarium of the Xishuangbanna Tropical Botanical Garden (HITBC), Chinese Academy of Sciences.
Edible species and their ethnic nomenclature
Wild edible Ficus species used by each ethnic group and their names
F. altissima Bl.
Da qing shu (lofty tree)
F. auriculata Lour. (F. roxburghii Wall.)
Xiang er rong, Mu gua rong (elephant ear fig or papaya fig)
F. callosa Willd.
Ying pi rong (hard skin fig)
F. hirta Vahl.
Wu zhi mao tao, Cu ye rong (five – finger hairy fig or rough leaf fig)
F. maclellandii King var. Rhododendrifolia Corner
Du juan ye rong (azalea leaf fig)
F. oligodon Miq.
Ping guo rong (apple fig)
nuoge zam biu
F. racemosa L. (F. glomerata Roxb.)
Ju guo rong, Ma lang guo (Cluster fig)
guole, mulu se*
F. semicordata B.-H. ex Sm.
Ji su guo (chicken crop fig)
mule lum biu
F. semicordata var. montana Amatya
Ji su guo (Small fruit variety of chicken crop fig)
mule zam biu
F. tikoua Bur.
Di ban teng, Di shi liu (ground pomegranate)
F. vasculosa Wall. ex Miq.
Tu mai rong, Shan tian cai (sweet mountain greens)
pàk dé gái
F. virens Ait.
Lv huang ge shu, Suan bao shu (sour buds tree)
F. virens Ait. var. sublanceolata (Miq.) Corner
Huang ge shu, Suan bao shu (sour buds tree)
In the Dai language, names of wild plants used as vegetables are often preceded by “pàk”. The words “liāng” (reddish), “xiū” (green), “háo” (white), “núi” (smaller) and “lōng” (bigger) are often added to discriminate the edible parts of similar fig plants. Some Ficus forms distinguished by local people are apparently not yet recognized taxonomically. Sometimes, the flavor is reflected in the plant name. For example, the Dai word for chicken, “gái.” is used in the Dai name for F. vasculosa, describing the chicken-soup like taste of the young leaves used in vegetable soup.
In the Hani language, “xibu” refers to the fig fruits borne in clusters on the trunk or thick leafless branches, while “xigu” refers to fruits borne in long and slender leafless braches from the trunk and near the ground. The words “ha” (bitter, here means inedible male figs), “ma” (bigger) and “qi” or “misi” (smaller) are often added after the formal name. The Hani word “niza” means a guest who stayed too long and can’t be driven away, and is used for strangling fig trees! Typical strangler species, such as F. virens, are disliked by the Hani even though they know the leaf buds are edible.
In the Jinuo language, “neme” refers to leaf buds with a sour taste and “adao” is the common name for leaves. In the Yao language, “nuoge” is a common name for edible plants with white latex. The words “lumu” and “zamu” refer to “bigger” and “smaller” fig fruits. “Biu” is a common name for edible fruits, and “den” means young leaves or leaf buds.
Collection and form of consumption
The growth habit, habitat, harvest season and consumption form of the Ficus found in the ethnic diet in Xishuangbanna, SW China
Growth habit (M/D)B
Intensive harvest period
Mode of consumptionC
Large trees (M)
Insides and margins of the thick forest in valleys or mountains, or grow as individuals in plains
February – March
Vegetables boiled with pork ribs (water blanching before cook, Hani), the stipules act as sour taste ingredients
Small trees (D)
Tropical or subtropical forests in moist valleys, or surroundings of farmland and village
January – April (young leaves), March – July (ripe female figs)
Young or ripe female figs, young leaves
Ripe figs are eaten raw or for making jelly beverage; Young figs are used as salad with condiments (Dai and Jinuo); Young leaves are used as vegetables boiled with spareribs (water blanching or rubbing with salt before cook); Young figs and young leaves are sold for cash income
Large trees (M)
Forests in basins or valleys in lower mountains
January – May
Vegetables cooked with tomato (the red kind need water blanching before cook, Dai, Jinuo); sold for cash income
Shrubs or small trees (D)
Slopes or margins of mountain forests or open fields near villages
August – October
Ripe female figs
Child snacks (Hani, Jinuo and Yao)
Plains or thin forests along river and stream sides
February – April
Fresh vegetables, or store up after quickly baked and dried for use out of season (Dai); Being sold for cash income
Small trees (D)
Forests in higher mountainous areas,
(Similar to FAU)
Ripe female figs, young leaves
Ripe figs are eaten raw or for making jelly beverage; Young leaves are used as vegetables boiled with spareribs (water blanching or rubbing with salt before cook); Young leaves are sold for cash income
Large trees (M)
Thin forests along river and stream sides, or valleys of lower mountains
March – May
As salad with condiments or as vegetables cooked with green moss (Dai, Hani, Jinuo)
Small trees (D)
Forest edge or thin forests in valleys, beside rivers and roads
Irregular (2 – 3 crops per year)
Ripe female figs
Ripe figs are eaten raw
Small trees (D)
Forest edge or road side
Irregular (3 crops per year)
Ripe female figs
Ripe figs are eaten raw
Prostrate woody vines (D)
Slopes of limestone mountain and grass land at higher elevations
June – September (ripe female figs)
Ripe female figs; whole plant
Ripe female figs are eaten raw; Whole plant is used for tea preparation (Yao)
Seasonal rain forests at lower elevations
January – June
Soup vegetables boiled with other wild greens or fried vegetables (Dai, Hani); Sold for cash income
Large trees (M)
Forests in valleys or lower mountains
January – April
As salad with condiments (Yao) or as vegetables boiled with pig trotter, the stipules give a sour taste; Being sold for cash income
Large trees (M)
Forests in valleys or lower mountains, or growing as individuals in plains
January – April
Vegetables boiled with pig trotter, the stipules give a sour taste (Dai)
The number and location of edible Ficus species kept by households in Manlun village in Xishuangbanna, SW China
Total number of plants
Percent of households with each speciesB
Number of plants kept by each household
F. virens var. sublanceolata.
Comparative evaluation of the usage intensities and preference
Comparison of the usage intensities of each edible Ficus species among different culture groups
+ + + +
+ + + +
+ + + +
+ + +
+ + + +
+ + + +
+ + + +
+ + +
+ + +
+ + +
+ + +
+ + +
+ + +
Comparison of the wild edible Ficusspecies uses among the four culture groups
The Dai, Hani, Jinuo and Yao represent four distinctly different cultural groups with different languages in XSBN. All four groups have accumulated extensive knowledge on using the female fruits (For dioecious fig taxa, only the female fruits are edible, they are seed producers and are larger in size, rich in nutrient and have a pleasant taste, while the male fruits function in breeding pollinator wasps are poor in nutrient and not palatable. For monoecious fig taxa, their fruits are the same and avoided by people because small size, poor taste and the presence of wasps), young leaves or leaf buds of Ficus species as food resources. The common consumed species across the four groups are F. auriculata, F. oligodon, F. semicordata, F. semicordata var. montana and F. virens. The wide consumption and sustained marketing of these species show that they are widely accepted food resources rather than a narrow cultural preference. All the species had fixed and usually food-related ethnic names, which indicates the long history of indigenous consumption.
Different cultural groups use the wild edible Ficus species to different extents, depending on their availability, the group’s dependence on forest resources, and cultural factors. Generally, the Dai people use them at the highest intensity because (i) they have been permanent inhabitants in XSBN for centuries, and have established homegardens for small-scale wild food or medicinal plant cultivation or domestication, and (ii) they live in basins and have better access to markets. The other three mountain culture groups have more recently established permanent living spaces in the region and do not have a culture of homegarden cultivation, depending more on forest collection. Our data show that the Hani people in Nanbang prefer fruits of F. auriculata and F. oligodon (“Xibu”). One of their orally transmitted legends tells that the first Hani people were born in a “Xibu” and survived by sucking the “milk” in it (the milky latex that characterizes figs). In addition, their ethnic epic records that “Xibu” is one of the first edible fruit plants they discovered in the forest during hunting activities. The villagers do not like strangling figs, such as F. alltissima and F. virens, since they often encounter huge strangling figs in the forest which have killed their host trees, and believe this is an omen of disaster. They therefore remove most of spontaneous seedlings around the village.
The significance of the cultivation and commercialization of the wild edible Ficusspecies
The possibility of a wild plant resource continuing to meet both subsistence and market demands largely depends upon sustainable harvest by appropriate management, and the domestication of wild resources is crucial for resource management . Domestication of common fig is thought to have started in the Mediterranean region in Early Neolithic period  and then spread worldwide. In XSBN, the Hani, Jinuo and Yao protect wild edible Ficus species while the Dai cultivate them. Except for the species traditionally cultivated in or around courtyards for ornamental and consumption purpose (such as F. auriculata), the cultivation of most Ficus species was triggered by the expansion of rubber plantations and resulting sharp decline in wild resource accessibility since the early 1980s. The indigenous villagers prefer to plant these species for their wide adaptability, high productivity, easy management and pleasant palatability. Ficus species are cultivated as tree fodder in many parts of the Himalaya region, with F. auriculata the most widely used species. Its fodder quality is far superior to paddy straw . In the Bamileke region of Cameroon, Ficus species are propagated by pole cuttings and are an important part of agrarian system management . These examples show that wild Ficus species undergo a gradient of management intensities from simple gathering, to nonselective incipient management, selective incipient management and occasionally ex situ cultivation . But whether or not the incipient domestication has occurred is unknown.
The commercialization of wild edible Ficus species in XSBN is an important source of cash income for indigenous villagers. The leaves, buds and fruits of several Ficus species are sold in local markets and the whole plants are sold as ornamental trees. In other regions, however, only in north and central Vietnam are the near-ripe peeled or unpeeled fruits and young leaves of elephant figs (F. auriculata Lour. or F. oligodon Miq.) on sale in the market, and in Papua New Guinea the young leaves of dinner-plate figs (F. dammaropsis Diels.) are commonly sold in highland markets . As Sawian et al.  reported, these products seldom appear to contribute a large share of a household’s total income generation, but are often important in bridging seasonal or other cash flow gaps.
The characteristics of the wild edible Ficusspecies in Xishuangbanna compared to those in other regions
The use of wild Ficus species as food is widespread in areas where the genus occurs, especially in the Himalaya region, which is floristically similar to our study area. For the 13 edible Ficus species in this study, the consumption of 6 species has either not been reported from elsewhere (F. altissima, F. callosa, F. maclellandii var. rhododendrifolia and F. vasculosa), or only from South China (F. tikoua and F. virens var. sublanceolata). Consumption of the other 7 species had been reported from Nepal [7, 27] and some were reported as eaten in India [9, 13, 28, 29], North Laos , Vietnam  or Pakistan [16, 30].
No other region appears to consume the diversity of wild Ficus species eaten in XSBN. Moreover, in the species consumed only in XSBN as well as those used most intensively, leaves and buds are the major parts consumed. These are concentrated sources of vitamin E, vitamin B1 (thiamin), vitamin B2 (riboflavin), protein and minerals , and are apparently also rich sources of naturally occurred antioxidants , suggesting that they may make a significant contribution to the health and well-being of the consumers. Antioxidant potential has also been demonstrated in some other Ficus leaf samples .
Our studies of wild, managed in situ and cultivated edible Ficus populations showed that their edible products are highly appreciated by the indigenous people of Xishuangbanna. We found that in both mountain villages, which have forest access but are far from markets, and basin villages, which have market access but no forests, figs are used as vegetables, fruits or beverages. Moreover, people in both situations invest effort in promoting the use of these species through artificial management. The Dai people who live in basin villages cultivate preferred species in and around their kitchen gardens and courtyards, encouraging intensive usage, with the possibility of artificial selection of plants with preferred characteristics. Further studies should be conducted to determine if these species are or can be ongoing incipient domestication. Finally, this study of a small geographic area suggests that the genus Ficus represents a largely untapped source of potential food resources for tropical people.
The authors are grateful to the National Natural Science Foundation of China (31400286) and the Ministry of Science and Technology of the People’s Republic of China (1zY213631271) for financial support. Professor Darong Yang identified the Ficus species and contributed valuable suggestions and encouragement. Professor Zhekun Zhou and Ping Zhang supported the research platform and gave instructive comments. Professor Richard Thomas Corlett and Dr. Pelin Kayaalp edited the English language thoroughly.
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